J.M.C. HUTCHINSON 1990. Stabilising selection in Weldon’s snails: a reappraisal. Heredity 64:113–120.

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Update

I overlooked another paper that applied a similar approach with scale number in lizards: Cook, L.M. 1979. Variation in the Madeiran lizard, Lacerta dugesii. Journal of Zoology 187: 327–340.

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